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It's puzzling that his nails came off even with the regular clippings. I don't know why they would have come off other than from some sort of trauma, getting jerked off by being stuck somewhere. As to the clumps of fur, it could be shedding time though it's a little late for that. It usually happens around August or Sept. Is it possibly a skin problem or mites? Take a look at the info on this page and see if you can identify some other symptom: http://www.bio.miami.edu/hare/furloss.html
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